Smilodectes gracilis (Notharctidae)



Smilodectes gracilis ist eine Primatenart innerhalb der Familie Notharctidae, die ab dem Paläogen (Eozän) im Ypresium lebte, das vor rund 56 Millionen Jahren begann und bis vor 47,8 Millionen Jahren andauerte.

Systematik

Daten zu den einzelnen Funden von Smilodectes gracilis
Sammlung Epoche, Alter Geologie, Formation Kommentar zur Sammlung
1375 No.8 Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger B-8
U.a. am Fundort ausgegraben: Microsyops elegans
Sammlung Epoche, Alter Geologie, Formation
B-11 Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger
Sammlung Epoche, Alter Geologie, Formation
Cottonwood Creek (Lower Bridger) Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger
Kommentar z. Taxonomie
"Bridger C is exposed along the upper course": Wood 1934
"Viverravus gracilis" Thorpe 1923a: level not stated

AMNH 19231 from B3 is either "Eotrigonias petersoni" or Hyrachyus: Wood 1927
U.a. am Fundort ausgegraben: Notharctus tenebrosus
Sammlung Epoche, Alter Geologie, Formation Kommentar zur Sammlung
Fault Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger B-1; Big Sandy; UW V-570001
Kommentar z. Taxonomie
due to errors and omissions in their list, I am unsure if the following "Bridger, Big Sandy" records of Black and Sutton 1984 pertain to Fault: Paramys excavatus Leptotomus parvus "Ischyrotomus oweni"
U.a. am Fundort ausgegraben: Omomys carteri
Sammlung Epoche, Alter Geologie, Formation
Grizzly Buttes Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger
Museum Kommentar z. Taxonomie
AMNH,USNM,YPM mostly from Bridger B2 according to Matthew 1909 and Osborn 1929
type of "Triacodon nanus" is from Grizzly Buttes: Thorpe 1923a
U.a. am Fundort ausgegraben: Microsyops elegans
Sammlung Epoche, Alter Geologie, Formation Kommentar zur Sammlung
Hawk Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger B-3
U.a. am Fundort ausgegraben: Omomys carteri
Sammlung Kommentar zum Fundort Epoche, Alter Geologie, Formation Kommentar zur Sammlung
Huerfano I "Sec. 14, 15, 16, T26S, R70W" Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Huerfano Huerfano Muddy Divide, Muddy Fork
Kommentar z. Taxonomie
Gardnerbuttean; "Most specimens are from below the white bed (unit 20 of measured section)"
U.a. am Fundort ausgegraben: Microsyops scottianus
Sammlung Epoche, Alter Geologie, Formation Kommentar zur Sammlung
Jean's Quarry Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger B-9
U.a. am Fundort ausgegraben: Microsyops elegans
Sammlung Epoche, Alter Geologie, Formation Kommentar zur Sammlung
Little White Butte Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger B-7
U.a. am Fundort ausgegraben: Microsyops elegans
Sammlung Epoche, Alter Geologie, Formation
Morrow Creek Locality 10 Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Green River
Kommentar z. Taxonomie
said to be in the "Morrow Creek Mbr.," which is actually the Laney Mbr. according to Gazin 1962
U.a. am Fundort ausgegraben: Omomys carteri
Sammlung Kommentar zum Fundort Epoche, Alter Geologie, Formation
North Fork of the Shoshone River (NF-4) Wapiti Valley Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Aycross
Kommentar z. Taxonomie
level is approximate
said to be Bridger B
U.a. am Fundort ausgegraben: Washakius insignis
Sammlung Kommentar zum Fundort Epoche, Alter Geologie, Formation
Picket Lake near Honeycomb Buttes, Great Divide Basin Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Green River
U.a. am Fundort ausgegraben: Cantius nunienus
Sammlung Kommentar zum Fundort Epoche, Alter Geologie, Formation Kommentar zur Sammlung
Powder Wash (Upper) 2 mi SE of Powder Springs Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Green River Powder Springs
Lithographie Kommentar z. Taxonomie
"a uniformly muddy, fine-grained sublitharenite" (Gunnell and Bartels 1999) sec 8 T 7S R 25E: Van Houten 1945
same as the "Green River" fauna of Burke 1935 and Kay 1957
according to Remy 1992 there is an Ar-Ar date of 47.2 Ma on the "curly tuff... between the S1 marker unit and the Magohany oil shale bed" (S1 unit is lower); because the fauna is from "82 m below" the shale and the date is interpreted by Remy as Uintan, I assume it is too young for use as a calibration point
"Class 1" reliability K-Ar date of 44.8 (= 46.0) +/- 0.9 Ma (Mauger 1977) for the Wavy Tuff is definitely too young for use as the tuff is 20 ft above the Mahogany Oil Shale; Wavy Tuff redated at 42.3 +/- 2.0 (FT on zircon) by Bryant et al. 1989
original list repeated almost without modification by Gunnell and Bartels 1999
Black and Sutton list the following: Thisbemys corrugatus "Leptotomus bridgerensis"
falls in the Parachute Creek Mbr. sensu Roehloer 1992
early Bridgerian
the record of Apatemys hendryi reported by Robinson 1966b is omitted without comment by K & S
U.a. am Fundort ausgegraben: Uintasorex parvulus
Sammlung Epoche, Alter Geologie, Formation Kommentar zur Sammlung
Tree Road Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger B-4
U.a. am Fundort ausgegraben: Omomys carteri
Sammlung Epoche, Alter Geologie, Formation
Twin Buttes (Lower Bridger) Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger
Kommentar z. Taxonomie
Twin Buttes records of unclear status: "Notharctus robustior" Robinson 1957: supposedly upper Bridger, but probably lower Bridger based on Gazin's discussion of Smilodectes; could be N. pugnax or perhaps N. tenebrosus based on chart Patriofelis ferox Wortman 1894: "Twin Buttes" Limnocyon verus Thorpe 1923a: level not stated Orohippus progressus type if from an unknown level at Twin Buttes according to Gazin but from "Hor. C" according to Granger 1908; see also Kitts 1957b, who implies that O. progressus and O. sylvaticus are present in Bridger C sensu Gazin Palaeosyops robustus Osborn 1929: from "C or "D" Telmatherium cultridens Osborn 1929: "C or D" Mesatirhinus megarhinus Osborn 1929: level not stated "Homogalax bridgerensis" Troxell 1922b: type, level not stated and Isectolophus latidens ranges throughout section according to Gazin Hyrachyus eximius Wood 1934: "Twin Buttes... hence Bridger C or D" and "South Twin Buttes"; referred specimen of "H. princeps"; all are possibly from Bridger B based on revised stratigraphy of Gazin Hyrachyus modestus and type of "Ephyrachyus cristalophus" of Wood 1934: both from "Bridger C3" and based on Wood's map it is not clear whether this is above or below the Sage Creek White Layer "agreeing in size with" Homacodon sp. Sinclair 1914: Bridger "C1"; specimen not mentioned by West 1984, and probably from Bridger B based on stratigraphy of Gazin 1976 "Lophiohyus alticeps" type is from an unknown level; see also Sinclair 1914
Sammlung Epoche, Alter Geologie, Formation Kommentar zur Sammlung
Wash Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger B-5
Kommentar z. Taxonomie
Korth 1984: "Microparamys sp. B" is either Mattimys or Janimus
U.a. am Fundort ausgegraben: Omomys carteri
Sammlung Kommentar zum Fundort Epoche, Alter Geologie, Formation Kommentar zur Sammlung
Bill’s Smilodectes BLM locality "in T 16N R 110W" (no section number or any other details given) Ypresian
zwischen 46.2 und 50.3 Millionen Jahren
Bridger DMNS loc. 885
Kommentar z. Stratigraphie Lithographie Museum
"approximately 1-m-thick horizon... about 4–5 m below Gazin’s (1965) Sugar White Layer" "slate green coloured mudstone" DMNH
U.a. am Fundort ausgegraben: Omomys carteri

Literatur

J. L. Wortman 1903, Studies of Eocene Mammalia in the Marsh Collection, Peabody Museum. Part II. Primates. Suborder Cheiromyoidea. The American Journal of Science, series 4. 16, p. 345 - 368
P. Robinson 1957, The species Notharctus from the Middle Eocene. Postilla. 28, p. 1 - 27
R. M. West 1973, Geology and mammalian paleontology of the New Fork-Big Sandy area, Sublette County, Wyoming. Fieldiana: Geology. 29, p. - 27
C. L. Gazin 1976, Mammalian Faunal Zones of the Bridger Middle Eocene. Smithsonian Contributions to Paleobiology. 26, p. 1 - 25
P. D. Gingerich 1979, Phylogeny of middle Eocene Adapidae (Mammalia, Primates) in North America: Smilodectes and Notharctus. Journal of Paleontology. 53:1, p. 153 - 163
L. Krishtalka, R. K. Stucky 1984, Middle Eocene marsupials (Mammalia) from northeastern Utah and the mammalian fauna from Powder Wash. Annals of Carnegie Museum. 53:2, p. 31 - 46
G. F. Gunnell, W. S. Bartels, P. D. Ginberich, V. Torres 1992, Wapiti Valley Faunas: Early and Middle Eocene Fossil Vertebrates from the North Fork of the Shoshone River, Park County, Wyoming. Contributions from the Museum of Paleontology, University of Michigan. 28:11, p. 247 - 287
R. K. Stucky, W. S. Bartels, P. D. Ginberich, V. Torres 1994, Personal communication regarding identifications of Eocene mammals. . :11, p. - 287
J. Alroy, W. S. Bartels, P. D. Ginberich, V. Torres 2002, Synonymies and reidentifications of North American fossil mammals. . :11, p. - 287
M. Spaulding, J. J. Flynn, R. K. Stucky 2010, A new basal Carnivoramorphan (Mammalia) from the ‘Bridger B’ (Black’s Fork member, Bridger Formation, Bridgerian Nalma, middle Eocene) of Wyoming, USA. Palaeontology. 53:4, p. 815 - 832, DOI: 10.1111/j.1475-4983.2010.00963.x